Role of Carotenoids on the Immune Response

Role of Carotenoids on the Immune Response There is growing evidence from in vitro and in vivo laboratory animal studies that ÃŽ ²-carotene can protect phagocytic cells from autoxidative damage, enhance T and B lymphocyte proliferative responses, stimulate effector T cell functions and enhance macrophage, cytotoxic T cell and natural killer cell tumoricidal capacities, as well as increase the production of certain interleukins. Because of their molecular composition, specifically their highly conjugated double-bond structure carotenoids such as beta-carotene and lutein serve as effective scavenger of the harmful unpaired electrons (eg. singlet oxygen, free radicals) produced by normal metabolic processes that damage body tissues (eg. via lipid peroxidation, Krinsky 1992).The cells of the immune system are particularly sensitive to oxidative stress and may benefit substantially from the free radical trapping ability of carotenoids, as the immune response itself produces reactive oxygen species that disrupt the intercellular si gnals sent via lipid-rich, membrane- bound receptors (Chew 1993). An enormous body of literature has accumulated over the past 20 year demonstrating the potency of carotenoids as immuno stimulants in humans and other mammals(reviewed in Hughes 2001, Krisnsky 2001). However, within the last decade, it has also been suggested that these carotenoid derived sexual colours may signal the health and condition of males of birds and fish of the specific antioxidant and immune stimulatory activity of carotenoids obtained from the diet and transported through the body before they are incorporated into the integument (Lozano 1994, von Schantz et.al; 1999; Moller et.al 2000). This issue continues to be debated on theoretical grounds (Hill 1999a ; Lozano2001), fueld by the idea that colorful birds and fishes obtain far more carotenoids in the diet than carotenoid deprived mammals, thus individuals may not be limited in the extent to which they can use carotenoids to boost their immune response. Early studies demonstrating the ability of dietary carotenoids to prevent infections have left open the possibility that the action of these carotenoids may be through their prior conversion to vitamin A. Subsequent studies to demonstrare the specific action of dietary carotenoids have used carotenoids without provitamin A activity such as lutein, canthaxanthin, lycopene and astaxanthin. In fact, these non provitamin A carotenoids were as active, and at times more mediated and humoral immune response in animals and humans. Results have similarly shown immuno enhancement by non provitamin A carotenoids, based either on the relative activity or on the type of immune response affected compared to ÃŽ ²-carotene Studies on the role of carotenoids on immune response have generally used several key immune function assays. These include Ig production,lymphoblastogenesis, lymphocyte cytotoxic activity, cytokine production, delayed type hypersensitivity. In the 1930s, Green and Mellanby found that infections of the ear, bladder, kidney and gut of vitamin A-deficient rats were prevented when the rats were fed with ÃŽ ²-carotene. The level of carotene needed to prevent these infections was higher than the amount required to stimulate growth. Clauson observed that in young children with severe ear infections, improvement was seen following an increased intake of dietary carotene. These two early reports suggest that the anti-infective effect of ÃŽ ²-carotene may be due to its pro-vitamin A activity. ÃŽ ²-carotene may also have an independent effects on immune responses, separate from its provitamin A activity. ÃŽ ²-carotene and other carotenoids with nine or more conjugated double bonds may enhance immune function by quenching singlet oxygen and other reactive oxygen species, including free radicals. ÃŽ ²-carotene and canthaxanthin together inhibited the loss of macrophage receptors following exposure to reactive oxygen intermediate (Gruner et. al., 1986). T helper cells are involved in helping B cells secrete specific antibodies in response to an antigenic challenge. Experiments were performed , in which laboratory animals were either fed ÃŽ ²-carotene or canthaxanthin to determine a carotenoid effect, separate from a provitamin A effect on T and B lymphocyte functions. Canthaxanthin has the same ability to quench singlet oxygen and free radicals as ÃŽ ²-carotene but cannot be converted to vitamin A in mammals. Specific immune responses of laboratory animals were similarly enhanced with diets containing canthaxanthin as with diets containing ÃŽ ²-carotene (Benedict and Shapiro 1986). Some new investigations indicate that carotenoid can play a very important role in enhancing immune responses, which can lead to the reduction of tumor growth. The immune system has three cell types capable of killing tumor cells. In a preliminary report, it was shown that human natural killer cells killed significantly more tumor cells when incubated with ÃŽ ²-carotene than human cells not exposed to ÃŽ ²-carotene. ÃŽ ±-carotene also enhanced tumor killing (Leslie and Dubey, 1982). Carotenoids, besides acting through the various mechanisms described earlier, can also influence immune function through their ability to regulate membrane fluidity, and gap junctional communication. All these actions are most likely interrelated in their modulation of an immune response. Carotenoids help to maintain the membrane receptors that are essential for immune function, and they may also be found to be important in the release of immune modulatory lipid molecules such as leukotrienes. Carotenoids may increase the tumoricidal activity of the cytotoxic T cells, macrophages and or natural killer cells by any of these mechanism of action. Carotenoid enhances many aspects of immune function which include T and B lymphocyte proliferation, induction of specific effector cells capable of killing tumor cells, and the secretion of factors required for the communication between immunologically competent cells. The action of carotenoids on immune response hangs in a delicate balance with the intra-and extra cellular milieu, the outcome of which depends not only on the type and concentration of the carotenoid but also on the cell type and animal species involved. Even though studies to date have provided evidence for a specific action of carotenoids much has yet to be done to truly understand their molecular action. CAROTENOID AS PRO-VITAMIN A SOURCE Biosynthesis or conversion of carotendoids into vitamin A: Conversion of carotenoids into vitamin A becomes complicated in fishes and aquatic animals, due to the presence of vitamin A2, which frequently predominated over vitamin A in animals living or spawning in freshwater.The presence of an unsubstituted ÃŽ ²-ionone ring in the caroteniod molecule supports for vitamin A activity. Both vitamin A1, and A2 can be formed in some fishes from common precurors, such as ÃŽ ²-carotene (Morton and Creed, 1939). Carotenoids containing at least one unsubstituted 3, 4 dehydro ÃŽ ²-ionone ring have been shown to act as specific provitamins A2 in chicks (Budowski, et.al 1963) and mice (Budowski and Gross, 1965).According to (Budoswki, et. al 1963) the dehydration of lutein to anhydrolutein (3,4- dehydro-3-hydroxy ÃŽ ²-carotene), a provitamin A2 might be of possible biological significance in this respect. There are two pathways for the conversion of carotenoids to vitamin A in mammals, central cleavage and eccentric cleavage. In 1960, Glover thoughtfully discussed various pathways by which carotenoids might be converted into vitamin A. He indicated that two primary oxidative reaction might occur, one at the central 15, 15’ double bond and the other at one or more of the other double bonds. The product of the central cleavage of ÃŽ ²-carotene would be two molecules of retinal, where as that of eccentric cleavage would be one long and one short ÃŽ ²-apo-carotenal e.g. cyclocitral and ÃŽ ²-apo-8’-carotenal, by the rupture of the 7’:8’ double bond. The enzyme, ÃŽ ²-carotenoid-15, 15’-dioxygenase which converts ÃŽ ²-carotene into two molecules of retinal requires molecular oxygen and is inhibited by sulfhydryl-binding and iron –binding reagents. Most pro vitamin A carotenoids,including the ÃŽ ² apo-carotenals,are cleaved to retinal by this enzyme. Thus, carotenoid dioxygenase with eccentric bond specificity has been indentified in mammals, the yield of ÃŽ ²Ã¢â‚¬â€œapo-carotenals from ÃŽ ²-carotene in vivo and in vitro is very low, and ÃŽ ²-apo-carotenals are formed nonbiologically from ÃŽ ²-carotene. Cyclic carotenes may be oxidized to hydroxylated and epoxide derivatives, converted to allenic and less saturated derivatives, or cleaved oxidatively to shorter products. Central Cleavage: In 1965, the cytosolic enzyme preparations of the intestine and liver converted ÃŽ ²-carotene to retinal were shown by two independent groups (Goodwin, et.al and Olson et. al 1965 ). The enzyme required molecular oxygen, yielded retinal as the sole identified product, was inhibited by ferrous-ion chelating agents and by sulfhydryl binding reagents (Olson, 1983). The main yield of retinal formed from the ÃŽ ²-carotene was 40-60%( Goodman and Huang, 1965) and (Olson, 1969). Since ÃŽ ²-carotene is unstable during isolation by TLC some losses occurred as a result of both biological as well as non biological oxidation during incubation, retinal was the major, if not the sole, biological product of the reaction. The retinal formed was satisfactorily characterized by several chemical and physio-chemical procedures. The enzyme, termed ÃŽ ²-carotenoid 15, 15’- dioxygenase was subsequently purified 20 to 70 fold from the intestines of several species and upon purification it became unstable. ÃŽ ²-carotenoid- 15,15’-dioxygenase cleaves many carotenoids, including several ÃŽ ²-apo-carotenals.There may be some quantative differences but the relative rates of cleavage are similar(Singh and Cama 1974). In general, relative to the aldehyde form, the ÃŽ ²-apo-carotenals are better substrates and the ÃŽ ²-apo-carotenic acids are poorer substrates for the enzyme. Retinal has clearly been identified as one product of the reaction. Eccentric Cleavage. ÃŽ ²-apo-carotenals are intermediate between carotenoids and vitamin A. In conversion of ÃŽ ²-carotene to vitamin A, an important observation was made in a recent study that individual ÃŽ ²-apo-carotenals are formed in small but significant amounts by incubating ÃŽ ²-carotene aerobically in the dark with shaking for an hour in the absence of an enzyme preparation. When compared, the reported rate of enzyme-catalyzed retinal formation is much higher.Despite these low yields, ÃŽ ²-apo-carotenals might still be formed biologically from B-carotene in mammals. But in future studies, the non-biological formation of oxidation products of carotenoids must be carefully assessed and the rate of the eccentric cleavage reaction, both in vivo and in vitro, must be related to the rate of formation of retinal by central cleavage. The metabolism of the ÃŽ ²-apo-carotenals has been addressed primarily in terms of their conversion to vitamin A and their oxidation to ÃŽ ²-apo-carotenoic acids. As analogous of retinal, however, they are expected to be metabolized in a similar way. Thus a significant part of them may well be reduced to ÃŽ ²-apo-carotenols (Glover, 1960) and (Sharma et .al 1977) and subsequently esterified to ÃŽ ²-apo carotenyl esters. Such compounds like retinyl ester and esterified lipids, should be transported primarily in the Chylomicra. Fish are much more versatile than mammals in meeting their needs for vitamin A (Olson, 1983). Fish are able to convert astaxanthin (3,3’-dihydroxy, 4,4’-diketo-ÃŽ ²-carotene), canthaxanthin (4,4 diketo-ÃŽ ²-Carotene) and isozeaxanthin (4,4’-dihydroxy ÃŽ ²-carotene) to ÃŽ ² carotene. They can also convert anhydrolutein derived from lutein, to dehydroretinol (Goswami ;1987). The oxidative metabolites of carotenoids work as essential molecules in a wide variety of living organisms. Carotenoids are converted to biologically active products such as abscisic acid, trisporic acid and retinoic acid in plants, fungi and animals respectively. Their formation is mediated by enzymes that catalyze cleavage reactions against specific double bonds of cartenoids. Moreover, carotenoids vulnerable to oxidation have the potential to be converted to biologically active compounds by chemical transformation. In mammals, ÃŽ ²-carotene 15,15’-dioxygenase catalyzes conversion of ÃŽ ²-carotene to retinal (Olson and Hayaishi, 1965) and (Goodman and Huang 1965). Although the cleavage enzyme plays a crucial role in vitamin A formation, its properties have not yet been fully revealed. The regulatory mechanism of the dioxygenase in particular remains to be clarified in terms of the nutrition of pro-vitamin A carotenoids. Dietary ÃŽ ²-carotene solubilized in mixed micelles wit h bile components and hydrolyzates of dietary lipids is absorbed in intestinal cells. Retinal formed from ÃŽ ²-carotene is further converted to retinyl ester by retinal reductase and lecithin retinol acyltransferase with the aid of cellular retinol binding protien type-II (CRBP-II), and then incorporated into chylomicron (Ong,1993). Thus, the cleavage of ÃŽ ²-carotene in intestinal cells is closely linked to lipid and retinoid metabolism, and the regulation of the cleavage enzyme activity should be considered in this context. Moreover, the cleavage enzyme is located in the intestinal cells, which are directly exposed to various food components. Action of dietary phytochemicals on the dioxygenase activity might affect the bioavailability of provitamin A carotenoid derived from fruits and vegetables. Carotenoids can be oxidized to a number of compounds by chemical transformation, in contrast to the specific cleavage of certain carotenoids by the enzyme, because of the high reactivity of conjugated double bonds to active oxygen species. The enzymatic conversion of ÃŽ ²-carotene to vitamin A, the non-enzymatic cleavage of lycopene, and the biological actions of the oxidation products of lycopene are described in Akihiko Nagao†s paper.

Does Free Trade Retard or Improve Development in Developing Countries E

Introduction The 19th century international system was a very significant era of the doctrine and practice of free trade. This key date in the 19th century was 1846, the year England repealed the corn law. The corn laws had protected British aristocrats and farmers from imported corn. Corn at this time was used to make bread. This dispute pitted industrialists and many workers against landowners and farmers. The industrialist wanted cheap corn so that the cost of bread would be lowered and they could reduce or reduce workers wages. This would give their products a competitive advantage in international markets over products from nations where the cost of food and wages were expensive. The workers in Britain also supported this law because they wanted to buy cheap bread. David Ricardo, a stock trader and member of parliament, articulated the liberal theory of comparative advantage which made intellectual case for free trade. Today this theory of comparative advantage has remained the bedrock of ar guments for economic integration and free trade. The broad purpose of this paper is to try and answer these questions; †¢ Does Free Trade Improve or Retard Development in Developing Countries †¢ Is Free Trade really Free. Free trade is the trade among two or more countries without any limitations imposed by the governments or other regulators; thus the free movement of goods and services across national frontiers (O’Brien and Williams, 2007 p.139). (Vander .W, 2005 p.24). It points to the liberalisation of the markets of nations through the elimination of tariffs, quotas and other form of restrictions to allow the free flow of products beyond national borders. Benefits of Free Trade Trade Openness. Free Trade has remained a fundam... ...the Barrel: Africa’s Oil Boom and the Poor. Available at http://www.crsprogramquality.org/storage/peacebuilding/Bottom%20of%20the%20barrel.pdf European Commission, (2012). Trade, growth and development; Tailoring trade and investment policy for those countries most in need [SEC (2012) 87 final]. O’Brien, R. and Williams, M. (eds.) (2007) Global Political Economy: Evolution and Dynamics. 2nd edition. Basingstoke: Palgrave. OECD, (2009). Journal on Development Trading Out of Poverty; How Aid for Trade Can Help. World Bank (2000) World Development Indicators 2000. Washington D.C: World Bank [Online] Available at http://data.worldbank.org/indicator Stop Free Trade : Available at http://www.stopftaa.org/is-free-trade-really-free-what-it-costs-a-country.html UNCTAD, (2004).The Least Developed Countries Report . UNCTAD/LDC/2004 (Overview), sales no.E.03.II.D.9.

Pro gay marriage Essay

How would you feel if someone tried to deny you the right to marriage? Does the government have the right to define marriage and decide who may marry? Marriage is one of the most common and powerful ways to show love and secure rights in that relationship. Marriage is many times taken for granted and even abused by many people. People tend to forget that the right to marry is not available to all people everywhere in the United States; some people have gone as far as proposing an amendment to the Constitution that denies same sex marriage. While churches can base their practice on their religious beliefs, civil rights, including the right to marry, should not be denied to anyone.How would you feel if someone tried to deny you the right to marriage? Does the government have the right to define marriage and decide who may marry? Marriage is one of the most common and powerful ways to show love and secure rights in that relationship. Marriage is many times taken for granted and even abu sed by many people. People tend to forget that the right to marry is not available to all people everywhere in the United States; some people have gone as far as proposing an amendment to the Constitution that denies same sex marriage. While churches can base their practice on their religious beliefs, civil rights, including the right to marry, should not be denied to anyone. In a country governed by the rule of law, the right to marry for homosexuals is inconsistently applied; and even when allowed, it is inconstantly recognized. Why should this be? The 1st amendment of the Constitution states that a person’s religious beliefs, or a lack of thereof, must be protected. Legislatures also cannot discriminate against marriages of the minority portions of the population; homosexual people fall into that category. There is also an amendment stating there is separation of church and state; therefore, religion cannot play a role in the civil rights issues of same sex marriage. Not t oo long ago, interracial marriage was frowned upon just like same sex marriage is today. (source 1) Imagine waking up one day to a world that was completely opposite from the world we knew when falling asleep, meaning gay people were now straight and all straight people were now gay. Do you think we would be willing to fight for our rights or have them denied; personally I would fight for my right to marry whom I choose, just as millions of Americans are currently doing. America is the land of the free, but we are not free to marry whomever we would like. We are equals in this world whether we are Black, White,  Hispanic, Indian, or any other race for that matter. Why should we not be equals based on sexual orientation? Churches must base their practices on their teachings; but basic civil rights under the law must be assured to all. No one in this nation is a second class citizen: marriage is a right under our legal guarantees of equality. Even though gay marriage should be legalized, some people have different beliefs. In most religions including Christianity, Islam, and Orthodox Judaism, same sex preferences may be a sin. Some decree that only homosexual activity is sinful. People believe in procreation to repopulate the world, but are against gay marriage and homosexuals adopting children to have the family they cannot create on their own. If this is truly the key issue against gay marriage, then it would only seem reasonable that only people w ho are fertile and want to procreate should be able to get married according to this bias. America has never really given gay marriage a chance, but seems to believe gay marriage will weaken this institution by leading to high divorce rates. Divorce rates are already high, even though most churches are not in favor of divorce. â€Å"One reason legislatures are denying gay marriage is because they are fearful that opening the door to allowing gay marriages will open the door to polygamy as well. This would also mean reopening cases about polygamy in the past which they are not willing to do.† (source 4) Others are fearful for children’s futures, and they fear being raised by two fathers or by two mothers may compromise children’s mental capacities and confuse their moral understandings. Marriage, based on true affection for another person, is not only a social intuition but an economic and safety measure as well. In places where gay marriages are banned, the gay couples are not able to have the same benefits as others. When filing for health care or insurance through a job, gay couples are not able to add each other on to their accounts. Most loved ones, such as a spouse or a child, in a straight marriage, can make life altering decisions in a hospital if need be. Since gay marriage is not legal, said person’s spouse is not recognized as the next of kin and care is delayed or wishes denied. (Source 2) California, Hawaii, New York, and the District of Columbia all have domestic partnership laws and civil partnerships, meaning it is a marriage without the religious factor. Civil partnerships also give couples the ability to have joint bank accounts, live in the same house, and pay bills together. Homosexual couples  are not entitled to this in most states. Ma rriage benefits should be available to all couples, no matter what. Parenthood is a benefit of marriage but gay couples cannot have children by themselves, without some type of help. They often look to adoption agencies. In some cases, gay couples are put on longer waiting lists or denied completely. There are other benefits to being married as well, such as tax breaks, a shared last name and adoption sharing as well as legal standing in society. It would seem that the solution to the question of same sex marriage would be easy; allow an individual the right to choose a partner and marry whom he or she wants. However, the issue in our society is intertwined in questions or religion, politics, ethnicity, and tradition. These factors affect every major issue in the nation today. Although I and many others support the right to marry, overcoming societal hurdles will not be easily accomplished. We must be heard on this issue, however long it takes to make the needed changes. It would seem that the solution to the question of same sex marriage would be easy; allow an individual the right to choose a partner and marry whom he or she wants. However, the issue in our society is intertwined in questions or religion, politics, ethnicity, and tradition. These factors affect every major issue in the nation today. Although I and many others support the right to marry, overcoming societal hurdles will not be easily accomplished. We must be heard on this issue, however long it takes to make the needed changes.

The Field Of Electrical Engineering - 1548 Words

The field of Electrical Engineering is the field into the future. Electrical Engineers learn electricity’s principles, interact with electrical properties, and imagine its application for the future. Without any end in sight, the field is vast and growing with several thousands of companies dominating the field today. Additionally, electrical engineering is proving to be a stable career choice as there is a growing market for electrical innovations, advancing technology to assist in the education of future engineers, and a high demand for workers. However, there is one large underlying problem for the entire field which surprisingly has nothing to do with the field itself: a lack of communication. In the age of the internet, millions of companies, engineers, and students still struggle to maintain modern knowledge of cutting edge technology, current inventions, and important discoveries due to the lack of common, free communication of these findings. 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